Climate, vegetation and fire history during the past 18,000 years, recorded in high altitude lacustrine sediments on the Sanetti Plateau, Bale Mountains (Ethiopia)

Abstract

Low-altitude lakes in eastern Africa have long been investigated and have provided valuable information about the Late Quaternary paleohydrological evolution, such as the African Humid Period. However, records often suffer from poor age control, resolution, and/or ambiguous proxy interpretation, and only little focus has been put on high-altitude regions despite their sensitivity to global, regional, and local climate change phenomena. Here we report on Last Glacial environmental fluctuations at about 4000 m asl on the Sanetti Plateau in the Bale Mountains (SE Ethiopia), based on biogeochemical and palynological analyses of laminated lacustrine sediments. After deglaciation at about 18 cal kyr BP, a steppe-like herb-rich grassland with maximum Chenopodiaceae/Amaranthaceae and Plantago existed. Between 16.6 and 15.7 cal kyr BP, conditions were dry with a desiccation layer at ~ 16.3 cal kyr BP, documenting a temporary phase of maximum aridity on the plateau. While that local event lasted for only a few decades, concentrations of various elements (e.g. Zr, HF, Nb, Nd, and Na) started to increase and reached a maximum at ~ 15.8–15.7 cal kyr BP. We interpret those elements to reflect allochthonous, aeolian dust input via dry northerly winds and increasingly arid conditions in the lowlands. We suggest an abrupt versus delayed response at high and low altitudes, respectively, in response to Northern Hemispheric cooling events (the Heinrich Event 1). The delayed response at low altitudes might be caused by slow negative vegetation and monsoon feedbacks that make the ecosystem somewhat resilient. At ~ 15.7 cal kyr BP, our record shows an abrupt onset of the African Humid Period, almost 1000 years before the onset of the Bølling–Allerød warming in the North-Atlantic region, and about 300 years earlier than in the Lake Tana region. Erica pollen increased significantly between 14.4 and 13.6 cal kyr BP in agreement with periodically wet and regionally warm conditions. Similarly, intense fire events, documented by increased black carbon, correlate with wet and warm environmental conditions that promote the growth of Erica shrubs. This allows to conclude that biomass and thus fuel availability is one important factor controlling fire events in the Bale Mountains.

Introduction

East African lakes have attracted scientific interest for decades. This is because global atmospheric circulation systems strongly influence this region and its hydrological dynamics (e.g. Thompson et al. 2002; Costa et al. 2014; Lamb et al. 2018). For instance, during the Last Glacial Maximum (LGM; 23–18 kyr BP) the decrease of tropical sea surface temperatures resulted in glaciation on East African Mountains (Mount Kenya, Kilimanjaro, Ruwenzori, and Ethiopian highlands) (Osmaston et al. 2005; Mark and Osmaston 2008). In addition, the southward shift of the Intertropical convergence zone (ITCZ) during ice rafting episodes throughout Heinrich event 1 (H1) are considered as a major cause for the dry event in northern and southeastern Africa around 16–17 cal kyr BP (Tierney et al. 2008; Marshall et al. 2007; Stager et al. 2011; Mohtadi et al. 2014). During the African humid period (AHP) maximum northern Hemisphere summer insolation shifted the rain belt associated with the ITCZ to the north (Bastian et al. 2021), generating increased rainfall across Northern Africa which turned the Saharan desert into green savanna (Gasse 2000). Palaeoclimatic studies suggest the southward extension of the AHP to eastern Africa (Tierney et al. 2008; Tierney and DeMenocal 2013; Costa et al. 2014).

Besides, lacustrine sediments (Tiercelin et al. 2008; Bittner et al. 2020; and many others), tropical glaciers (Gasse 2000; Thompson et al. 2002), peat bogs (Bonnefille and Mohammed 1994; Brown et al. 2007) and marine sediments (Camuera et al. 2021; Tierney and deMenocal 2013) have been serving as potential archives for the reconstruction of paleoenvironmental fluctuations in eastern Africa. However, the regional, topographic, and climate complexities, together with dating inaccuracy and lack of unambiguous proxies, lead to differences in the reconstructed timing of the onset and termination of major climatic events such as H1 and AHP across sites (Lamb et al. 2007; Tierney et al. 2011; Costa et al. 2014; Bastian et al. 2021). Moreover, the impact of these drastic climatic events on the migration of humans and on vegetation dynamics is yet not fully understood.

The highlands of the Bale Mountains in southeastern Ethiopia are promising sites for studying paleoenvironmental fluctuations. Their unique exposition to main atmospheric circulation systems (SW monsoon, SE monsoon, NE trades, and NW disturbances) (Miehe and Miehe 1994; Hillman 1986; Camberlin 2018) contribute to their climate sensitivity. In addition, the special geomorphological features, characterized by high altitudes up to 4377 m above sea level (asl), by moraines and small glacial depressions, support the reconstruction of the landscape evolution (Nicholson 2000). During MIS 3, the Bale Mountains were one of the most extensively glaciated mountains in Ethiopia (Messerli 1980; Osmaston et al. 2005; Groos et al. 2021). Groos et al. (2021) suggest that about 265 km² of the Bale Mountains were ice-covered between 42 to 28 kyr BP, well before the onset of the global LGM. This caused a temperature decrease and a downward shift of Afroalpine vegetation. Umer et al. (2007) and Ossendorf et al. (2019) suggested that the glaciers retreated from the Bale Mountains at ~ 15.9–16.7 cal kyr BP. Geochemical and pollen results from Lake Garba Guracha, in a north-exposed, deeply incised trough valley at 3950 m asl, and from peat deposits on the Sanetti Plateau, inform about Late Pleistocene-Holocene climate fluctuations (Tiercelin et al. 2008; Bittner et al. 2020) and respective vegetation changes (Umer et al. 2007; Kuzmicheva et al. 2013, 2014; Gil-Romera et al. 2019). These studies suggest that the warm and humid conditions during the early Holocene led to the expansion of Erica vegetation on the Sanetti Plateau, and dry climate and natural and/or human-induced fires might be major causes for the current patches of Erica on the Plateau. A study by Ossendorf et al., (2019), revealed human settlements in high elevation (3469 m asl) already during 47–31 cal kyr BP. Gil-Romera et al. (2019) reported that fire played a major role in determining the ecological dynamics of the Erica vegetation on the Bale Mountains. However, the influence of humans on the vegetation remains unclear. Open questions are: (1) Did the high altitudes of the Sanetti Plateau experience similar climatic fluctuations in intensity and timing as the lower regions in northern and eastern Africa during the Late Glacial? (2) Did the highlands of the Sanetti Plateau deglaciate at the same time as the north exposed trough valleys? (3) When and to what degree did Erica occupy the high altitudes of the Sanetti Plateau, where at present only patches can be found? (4) Are these patches of Erica relics documenting climate deterioration or human-induced fire?

In order to address these questions, and to better understand the paleoecological evolution of these high altitudes above the upper timberline, we analysed laminated lake sediments in high resolution, deposited in a glacial depression on the Plateau in about 4000 m asl using a multi-proxy approach. The depression has no noteworthy outflow and its catchment is not densely vegetated, thus being highly sensitive to local and regional environmental fluctuations.

Site information and methods/experimental

Study area

The Bale Mountains covering an area of 2200 km² include the Sanetti Plateau, which is the largest alpine ecosystem in Africa (Fig. 1). The area is widely perceived as natural (Miehe and Miehe 1994; Kidane et al. 2012). The Plateau (Fig. 1) extends between ca. 3800 to 4100 m asl, surrounded by the peaks of Tullu Konteh (4050 m asl) and Tullu Dimtu (4377 m asl), the second-highest peak of the country (Hillman 1988; Messerli and Winiger 1992). Erratic boulders, moraines, small lakes present on the plateau and in trough valleys are clear indicators of former glaciations (Osmaston et al. 2005). The parent rocks comprise Oligo-Miocene basalts and Quaternary rhyolites with trachytes, which weather to brown or brownish-black silty loams (Billi 2015). Muddy Gleysols are developed in depressions (Yimer et al. 2006).

Fig. 1

Location map of the study area. A Location of Ethiopia in Africa and regional location of the Bale Mountains National Park (BMNP) and Lake Tana. B Location of B4 depression on the Sanetti Plateau and location of Lake Garba Guracha. C Photo of B4 depression

Due to the annual migration of the ITCZ between 10° North and South, the climate of the Bale Mountains is characterized by a pronounced rainfall seasonality with a short dry (November to February) and a long rainy season (March to October) (Levin et al. 2009; Costa et al. 2014). The rainfall pattern is bimodal, with a peak from July to October, followed by a second peak from March to June (Kidane et al. 2012). The rain-bearing air masses derive from the Indian Ocean and the Atlantic Ocean via the Congo Basin (Gasse 2000; Tierney et al. 2011; Costa et al. 2014; Lemma et al. 2020), whereas northerly winds dominate during the dry season. In Dinsho (3170 m asl), the mean maximum temperature is 11.8 °C while the mean minimum temperature ranges from 0.6 to 10 °C, with frequent frost occurring in the high mountain areas during winter season (November to January) (Hillman 1986; Tiercelin et al. 2008). The highest precipitation and humidity occurs in the southern part of the mountains with 1000–1500 mm/a, while the northern part exhibits annual rainfall ranging between 800 and 1000 mm/a (Miehe and Miehe 1994; Umer et al. 2007).

The south to north rainfall gradient and the altitudinal temperature gradient stratify the vegetation into the Afromontane forest, the Ericaceous belt, and Afroalpine zone (Hedberg 1951; Friis 1986). The Afromontane forest spans from ~ 1450 to 3200 m asl in southern exposition, and from ~ 2800 to 3300 m asl along northern slopes. The southern declivity comprises Podocarpus gracilior associated with Syzygium guineense and Aningeria adolfi-friederici, whereas the northern slopes are mainly dominated by Juniperus procera, Hagenia abyssinica, and Hypericum revolutum (Friis 1986; Bussmann 1997). The Ericaceous belt covers ~ 90,000 ha between ca. 3200 and 3800 m asl and is dominated by Erica arborea L. and Erica trimera (Engl.) (Hailemariam et al. 2016). While the lower boundary of the Ericaceous belt (3300–3500 m asl) is covered with Erica-dominated Hagenia-Hypericum forests, the central part comprises monotonous Erica trimera stands, which continues to the upper limit (ca. 3800 m asl) of the boundary. The Afroalpine vegetation above 3800 m asl is open and rich in Tussock grasses and mainly dominated by Helichrysum splendidum-Alchemilla haumannii dwarf-scrubs and Giant Lobelia (L. rhynchopetalum) (Yineger et al. 2008), accompanied by patches of Erica, growing between big boulders along steep slopes (Miehe and Miehe 1994).

Sampling

We investigated a depression, located in an extended plain on the Sanetti Plateau, above the upper limit of the Erica belt at 3970 m asl (Fig. 1; 6° 53.3433′ S and 39° 54.5217′ E). A pit was dug down to the glacial boulders in a depth of 255 cm to prepare a profile named “B4” (Fig. 2). Subsequently, sediments were described (color, texture, structure, ect.) and sampled every 2 cm from 55 to 255 cm, representing humic-rich lacustrine, partly laminated material, and every 5 to 10 cm from the brownish upper part, rich in reddish mottles. Samples were air-dried and stored in plastic bags. In addition, 10 sediment samples were collected from the pit for radiocarbon dating. The air-dried samples were analyzed for elemental and mineral compositions, electrical conductivity (EC), pH (H₂O), stable C isotopes, alkanes, pollen, and black carbon (BC). Plant leaves (Alchemilla sp., Helichrysum sp., Pennisetum sp., Potamogeton thunbergii, and Ranunculus trichophyllus) and algae (Pediastrum sp.) were collected from the surrounding area to compare the stable isotopes composition of the sediment organic matter with its potential parent material.

Fig. 2

Stratigraphy and photo of the studied profile. A Photo of the B4 profile; the red pocketknife (10 cm) serves as scale. B Stratigraphy of the sediments in profile B4, divided in units 1–3, based on morphological features and calibrated radiocarbon ages (see Fig. 3; Table 1; YD = Younger Dryas, H1 = Heinrich Event 1)

Radiocarbon analysis

For the establishment of a reliable chronology, radiocarbon dates were obtained by dating alkali-soluble organic matter extracted from carbonate-free bulk samples using accelerated mass spectrometry (AMS) in the Radiocarbon Laboratory of the University Erlangen, Germany (Table 1). We established a Bayesian age-depth model using the IntCal13 calibration curve implemented in the package Bacon (Blaauw and Christen 2011) in R software (R Core Team 2013).

Table 1 Radiocarbon ages using Accelerated Mass Spectrometry of alkali soluble organic matter; calibrated according to Blaauw and Christen (2011)

Sediment analyses

Physical properties

Texture analyses were performed quantitatively in the laboratory on 52 air-dried and 2 mm sieved samples. Clay (< 6.3 μm) and silt (6.3–63 μm) fractions were quantified with a Mastersizer S (Malvern Instruments) after treating the samples with H₂O₂ and HCl. The sand fraction (63–2000 μm) was determined by wet sieving. We are aware that the fraction < 6.3 μm used in this study does not quantitatively correspond to the clay fraction defined by the pipette method as the fraction < 2 μm is underestimated by the Mastersizer (Antoine et al. 2009).

Inorganic geochemistry

For X-Ray Fluorescence (XRF) analyses, ground aliquots of 103 samples were dried at 105 °C. Major and minor elemental composition was determined using a Philips 2404 X-Ray Fluorescence Spectrometer. Given that the samples were carbonate-free, results were corrected for soil organic matter and water contents according to Eq. (1)

$$\% X_{{{\text{corrected}}}} = \% X_{{{\text{measured}}}} \times \left[ {\frac{100}{{100 - \%{\text{GV}}_{1000} - \%{\text{H}}_{2} {\text{O}}}}} \right]$$

(1)

X is the content of an element in percent and %GV₁₀₀₀ is the mass loss upon ignition at 1000 °C.

In order to evaluate the intensity of chemical weathering, the chemical proxy of alteration (CPA) was calculated according to Buggle et al. (2011) (Eq. 2).

$$\mathrm{CPA}=\left[\frac{{\mathrm{Al}}_{2}{\mathrm{O}}_{3}}{{\mathrm{Al}}_{2}{\mathrm{O}}_{3}+{\mathrm{Na}}_{2}\mathrm{O}}\right]\times 100$$

(2)

Qualitative mineral identification was carried out on bulk samples (n = 100) using X-ray diffraction. The powdered samples were packed into aluminum sample holders and scanned in a Philips PW 1710 diffractometer from 3 to 80°2^θ with Cu Kα radiation generated at 50 kV and 30 mA, at 0.02°2θ step size and 2.5 s step time. Phases identified in the samples are quartz, feldspar, pyroxene (probably mainly augite), clay minerals, hematite, pyrite, gypsum, and amorphous silica. No special clay mineral analysis was performed, yet characteristic peaks of clay minerals between ~ 6–12.5 and ~ 60–62°2^θ show a wide range of clay minerals, including smectite, chlorite, illite, kaolinite and mixed-layer minerals.

Organic geochemistry

Total organic carbon (TOC) and total nitrogen (N) of homogenized samples (n = 103) were quantified by dry combustion on a Vario EL elemental analyser (Elementar, Langenselbold, Germany). In order to complement the organic matter source identification and preservation status, n-alkanes and hydrogen index (HI) were analysed in 34 samples (Talbot and Livingstone 1989). n-Alkanes were extracted by soxhlet extraction using dichloromethane (DCM) and methanol (1:1) as solvents for 24 h (Zech and Glaser 2008). After adding 50 µl of 5α androstane as an internal standard, the excess solvent was removed by rotary evaporation and transferred to aminopropyl columns. Subsequently, n-alkanes were eluted with 3 ml of hexane DCM/MeOH (1:1). Finally, n-alkanes were quantified using gas chromatography (SHIMADZU, GC-2010, Kyoto Japan) equipped with an SPB-5 columns of 30 m length, 0.25 mm ID, 0.25 µm film thickness, and detected using a flame ionization detector (FID). Here, we used the n-alkane ratio Paq (C₂₃ + C₂₅)/ (C₂₃ + C₂₅ + C₂₉ + C₃₁) to identify the source of organic matter. HI was analysed by pyrolysis of dried (40 °C) and fine powdered bulk sediments (Talbot and Livingstone 1989).

We measured the natural abundance of δ¹³C (n = 101) by dry combustion of a 40 mg homogenized aliquot with a Fision 1108 elemental analyser coupled to a Delta S gas isotope ratio mass spectrometer (EL-IRMS) with a Conflow III interface (Thermo Finnigan MAT, Bremen, Germany). Sucrose (ANU, IAEA, Vienna, Austria) and CaCO₃ (NBS 19, TS limestone) were used as calibration standards for δ¹³C. The precision of δ¹³C measurement was 0.2‰. In addition to bulk stable isotopes, we measured compound-specific carbon isotopes (δ¹³C) of long chain terrestrial n-alkanes C₂₇, C₂₉ and C₃₁ to complement on the paleovegetation reconstruction (C3 vs C4). Compound-specific δ¹³C analyses of long chain n-alkanes (C₂₇, C₂₉ and C₃₁) were carried out and measured as described above.

Black carbon (BC) refers to polycondensed aromatic moieties formed during thermochemical combustion. While charcoal is usually detected visually using a microscope, BC is usually detected through chemical extraction, which enables to quantify the complex signature of pyrogenic carbon more precisely, especially when charcoal is not visible anymore. Thus, BC allows the reconstruction of fire events even if soot or char are not visible. Moreover, its polycyclic aromatic structure resists chemical and biological degradation in soil (Glaser et al. 1998; Wang and Li 2007; Wiedemeier et al. 2015) and persists for a long period of time in soil and sediments (Kuzyakov et al. 2014). We analysed BC (n = 72) using benzene polycarboxylic acids (BPCAs) as molecular markers following Glaser et al. (1998) and modified according to Brodowski et al. (2005). Five hundred mg of each sample was hydrolyzed with 10 ml 4 M TFA for 4 h at 105 °C. The hydrolyzed samples were filtrated on glass fiber filters and rinsed several times with de-ionized water to remove polyvalent cations. Subsequently, the samples were digested with 65% nitric acid for 8 h at 170 °C in a high-pressure digestion apparatus. The solution was passed through Dowex 50 W resin columns (200 to 400 meshes) to remove polyvalent cations. After derivatization, BPCAs were separated using GC and detected using FID with an injection temperature of 300 °C.

Palynomorph analysis

For palynological analyses, samples of 1 ml were prepared using standard procedures, including Lycopodium spores as exotic markers, acetolysis, HF treatment, and ultrasonic sieving (5 µm), and storage in glycerol (Stockmarr 1971). Microscopic analyses took place under 400 × magnifications, backed by oil immersion (1250 ×). For pollen identification, we used an existing reference collection of ~ 5000 slides (in Goettingen) and relevant literature (Gosling et al. 2013; Schüler and Hemp 2016). The nomenclature of the common types follows Beug (2004). Detailed analyses were carried out on 38 samples from the lower part of the profile (251–69 cm) with pollen counting sums of about 250 to 300 per sample. Pollen influx (grains cm⁻¹ a⁻¹) was calculated using exotic Lycopodium spore markers. To estimate humidity conditions, we calculated the A/C (Artemisia/Chenopodiaceae) ratio (Van Campo and Gasse 1993; Herzschuh 2007; Li et al. 2010) by dividing the number of Artemisia by that of Chenopodiaceae/Amaranthaceae. Poaceae pollen were size differentiated between small (< 37 µm) and large (> 37 µm) diameter grains. Counts of the larger grains allow an approximate estimate of C4 grasses, which produce larger pollen grains than C3 Poaceae (Jan et al. 2015). Diagrams were plotted with the software C2 (Juggins 2007).

Results

Age-depth model and sedimentation rates

B4 profile records sedimentation during the last 18.2 cal kyr BP. Our age-depth model (Fig. 3) illustrates that the sedimentation rate was 0.05 and 0.01 cm per year between 18.2 and 13.6 cal kyr BP and from 10.5 cal kyr BP to present, respectively. Between 13.6 and 10.5 cal kyr BP, a sedimentation hiatus is indicated.

Fig. 3

Bayesian age-depth model for profile B4 according to Blaauw and Christen (2011). Top left: The log-likelihood of the model fit for the saved iterations of the model, top middle: prior and posterior distribution of accumulation rate, and top right: prior and posterior distribution of the autocorrelation in accumulation rates (memory) and the time gap (hiatus size) at the end. Bottom panel shows the calibrated ¹⁴C dates (transparent blue) and the age-depth model (darker greys) indicate more likely calendar ages; grey stippled lines show 95% confidence intervals; red curve shows single ‘best’ model based on the mean age for each depth

Lithostratigraphy and interpretation of the depositional environment

The B4 sediments can be stratified into three units (Fig. 2). The lowermost unit 1 from 255 to 175 cm (18.2–16.6 cal kyr BP) is a dark-grey, laminated, silty clay with low sand content. It was deposited above a thin greyish sandy silt layer and basalt boulders. Unit 2, from 175 to 71 cm depth, is a grey, partly laminated silty clay, deposited between 16.6–13.6 cal kyr BP. A stiff sandy-silty light layer, preserved between 167 and 160 cm, likely documents intensive desiccation of the B4 depression at about 16.3 cal kyr BP. From 120 to110 cm (~ 15.4–15.2 cal kyr BP) plant root relics occur displaying swampy, shallow-water conditions. The upper lithostratigraphic unit 3 is about 70 cm thick, deposited during the last 10 cal kyr BP. It comprises light brown, weakly clayey, sandy silt with red mottles and bleached aggregate surfaces, showing water-logging during the rainy season. Unit 3 is strongly influenced by gleysation, which makes paleoenvironmental interpretations and regional comparisons ambiguous. Therefore, in the following, we present geochemical and palynological results only from units 1 and 2 in detail.

Grain size and geochemical results

Unit 1: 255–175 cm (18.2–16.6 cal kyr BP)

According to Fig. 4 and Additional file 1, clay and silt contents are high (partly up to 60% and 40%, respectively) and weakly fluctuating, whereas the sand contents are very low. TOC values, being low at the bottom of the core, constantly increase from ~ 0.7 to ~ 7% in 239 cm; then high values (~ 7%) persist until the top of unit 1. Low TOC/N values at the bottom of the profile increase to an average value < 12 (Fig. 4). HI values range between 107 and 266, with low values at the bottom of the profile.

Fig. 4

Analytical properties of the B4 sediments, with depth profiles of the sand fraction (63–2000 μm), total organic carbon (TOC), organic carbon-to-nitrogen ratio (TOC/N), Hydrogen Index (HI), δ¹³C values, Black Carbon (BC) and Paq. The light orange color shows dryer events while orange color indicates desiccation layers developed during Heinrich Event 1(H1). Blue color indicates the African Humid Period (AHP) and other humid episodes while dark green color indicates the event characterized as humid and arid according to pollen and geochemical results, respectively

Our results show an average δ¹³C value of − 14.3‰ with a slight decrease from the bottom to the top of unit 1. Analyses of water plants (Potamogeton thunbergii, Ranunculus trichophyllus) and algae (Pediastrum), sampled from nearby shallow lakes, resulted in δ¹³C values of − 18.6, − 15.5, and − 17.3‰, respectively. Modern terrestrial plants growing around the B4 depression are characterized by δ¹³C values between − 30.1‰ (Helichrysum argyaranthum) and − 26.9‰ (Alchemilla fischeri). Compound-specific analysis of long chain n-alkanes C₂₇, C₂₉, and C₃₁ revealed δ¹³C values of − 27, − 25.6, and − 25.1‰, respectively. Paq values increase from 0.27 at 244.5 cm to 0.44 at 175 cm. Despite high value at the bottom of the profile, BC values were very low in this unit (around 10 g kg⁻¹ TOC) increasing between 195 and 175 cm (~ 16.9–16.6 cal kyr BP) to around 30 g kg⁻¹ TOC.

Lithogenic elements such as Zr, Hf, Nb, and Na₂O show minimum values except for a slight increase at 210 cm and ~ 190 cm (~ 17.2 and ~ 17.0 cal kyr BP) (Fig. 5). Besides primary minerals such as quartz, feldspar, and pyroxene, unit 1 sediments contain traces of pyrite and gypsum. CPA values do not show fluctuation except a slight decrease at 17.2 and ~ 16.9 cal kyr BP.

Fig. 5

XRF results of B4 sediments. Depth profiles of Zirconium (Zr), Sodium oxide (Na₂O), Potassium oxide (K₂O), Hafnium (Hf), Niobium (Nb), Zirconium/Rubidium (Zi/Rb), Zirconium/Silica oxide (Zr/SiO₂), Zirconium/Titanium oxide (Zr/TiO₂) and of the chemical proxy of alteration (CPA = Al₂O₃/(Al₂O₃ + Na₂O), inversed scale. The light orange color shows dryer events while orange color indicate desiccation layers developed during Heinrich Event 1(H1) and the hiatus. Blue color indicates the African Humid Period (AHP) and other humid episodes while dark green color indicates the event characterized as humid and arid according to pollen and geochemical results, respectively

Unit 2: 175–71 cm (16.6–13.6 cal kyr BP)

Unit 2 starts at 175 cm depth with slightly increasing values of the lithogenic proxies (Fig. 5). Most striking is a pronounced sand maximum and a TOC minimum in ~ 160 cm, corresponding to ~ 16.3 cal kyr BP (Fig. 4). Strongly increasing values of the lithogenic compounds such as Zr, Na₂O, K₂O, Hf, Nd, and Nb and minimum CPA values are recorded between 160 and 135 cm (Fig. 5). Noteworthy is that these elements do not show the drastic excursion of TOC and sand in ~ 160 cm depth. Moreover, strong positive correlations (R = ~ 0.8–0.9; Additional file 2) are recorded between Na₂O, K₂O, Zr, Hf, Nb, and Nd in this unit. This constellation changed abruptly at ~ 15.7 cal kyr BP, documented by the abrupt decrease of most lithogenic elements and their ratios. However, some proxies such as sand content, TOC, HI, and δ¹³C do not show such an abrupt change except Paq values, which decrease between 140 and 120 cm (Fig. 4).

BC which was partly very low in unit 1, increased between 15.7 and 15.2 cal kyr BP (Fig. 4). The uppermost part of unit 2, (deposited between ~ 14.4 and 13.6 cal kyr BP (90–71 cm) is characterized by a distinct decrease in TOC, HI, varying sand contents, absence of pyrite, and higher contents of terrestrial n-alkanes and BC (Fig. 4).

Palynological results

The pollen record is divided into five local pollen zones (LPZ 1–5; Figs. 6, 7) based on changes in the pollen spectra. No pollen was detected in unit 3 due to pedogenetic processes that disturbed pollen preservation. About 75 pollen taxa have been identified in the record. The green algae Botryococcus and Pediastrum attributed to the intermittent presence of a water body in the B4 depression, thus their palynological record reflects local conditions.

Fig. 6

Percentage diagram of selected pollen taxa referring to the counting sum of pollen grains of a sample and fivefold exaggeration

Fig. 7

Pollen influx diagram of selected taxa and algae from B4 depression, showing pollen accumulation rate in grains cm⁻¹ year⁻¹

Poaceae < 37 µm dominate the pollen spectra with values around 35%. Other relatively common non-arboreal taxa are Artemisia, Chenopodiaceae/Amaranthaceae, Poaceae > 37 µm, and Plantago. The sum of arboreal pollen (AP) makes up around 15% in LPZ 1–4 (from 250.5 to 90 cm; 18.1–14.4 cal kyr BP) and increased to 38% during LPZ 5 (from 90 to 70 cm; 14.3–13.6 cal kyr BP). The green algae Botryococcus is most common in the lowest part of the record. Pediastrum increases with parallel decreases of Botryococcus. Pollen of modern water plants, Potamogeton thunbergii and Ranunculus trichophyllus are not present in the fossil pollen record.

LPZ 1 (250.5–204 cm; 18.1–17.2 cal kyr BP) is characterized by Botryococcus maxima, which abruptly decreases at 17.3 cal kyr BP and is replaced by high amounts of Pediastrum pollen (Fig. 7). Moreover, high percentages of Poaceae, herbs (non-arboreal pollen, NAP), Artemisia, and Chenopodiaceae/Amaranthaceae are recorded in LPZ1. Plantain (Plantago) has been very common. Arboreal pollen of Ericaceae, Podocarpus, Myrica, Juniperus, and Olea are found from the beginning of the archive.

LPZ 2 (204–194 cm; 17.2–16.9 BP) records an increase in Chenopodiaceae/Amaranthaceae and Plantago and strongly reduced occurrence of Artemisia and Pediastrum. Similarly, pollen percentage and pollen influx of Hagenia, Podocarpus, Myrica, Juniperus, and Olea show low values between 17.2 and 16.9 cal kyr BP.

LPZ 3 (194–110 cm, 16.9–15.2 cal kyr BP) starts with a subsequent increase of Podocarpus, Juniperus, and Olea from 194 to 177 cm (16.9–16.6 cal kyr BP) and a higher presence of Pediastrum. The afro-alpine element, Alchemilla, which is highly underrepresented in the pollen record, is sporadically present from now on. The influx of Erica, Pediastrum, Hagenia, and Botryococcus pollen decrease between 165 and 155 cm (16.3 and 16.1 cal kyr BP), while Chenopodiaceae/Amaranthaceae pollen increase. Subsequently, higher Pediastrum values, more frequent fern spores, and Sparganium-type are recorded between 155 and 135 cm (16.1 and 15.7 cal kyr BP). A lower occurrence of Erica pollen and the montane forest tree Hagenia recorded from 15.4 to 15.2 kyr BP, while absolute values of montane forest taxa like Podocarpus and Juniperus increase. By contrast, Pediastrum, Plantago-type, and Grass pollen influx show high values.

LPZ 4 (110–90 cm, 15.2–14.4 cal kyr BP) records the most abundant Ericaceae and Podocarpus. Similarly, the sum of all arboreal pollen (AP) increases but does not exceed their former values. The same holds true for Artemisia. In contrast, the ratio of Chenopodiaceae/Amaranthaceae falls as low as never before. In consequence, the A/C ratio increases strongly.

LPZ 5 (90–71 cm, 14.4–13.6 cal kyr BP) is marked by a strong expansion of Erica and Podocarpus. The previously significant elements Chenopodiaceae/Amaranthaceae and Plantago disappeared nearly completely, and Artemisia reduced strongly. Besides Poaceae, a wide range of Asteraceae (Senecio, Matricaria-type, Cichorioideae), which was present since the beginning of the record, became the most important vegetation constituents.

Discussion

Radiocarbon dating

Our radiocarbon ages show that depression B4 became ice-free at ~ 18 cal kyr BP and thus records the climate, vegetation, and fire history from the end of the LGM to the Late Holocene. The timing of deglaciation (18.2 cal kyr BP) seems reliable, as the sediments are free of carbonates.

Organic geochemistry

Low TOC contents at the bottom of the profile (Fig. 4) mark cold conditions with low biomass production in, and around, the lake. TOC/N ratios are used to distinguish between terrestrial and aquatic carbon sources. Organic matter derived from algae usually shows TOC/N values between 4 and 10, while vascular land plants and terrestrial soil organic matter may have TOC/N ratios > 20 (Meyers 1994) and ~ 15, respectively (Mekonnen et al. 2019). However, the use of TOC/N ratios as organic matter source indicators is highly constrained by degradation and mineralization, which results in low TOC/N values. TOC/N values of the B4 depression range between 3.5 and 12.3, suggesting mainly primary lacustrine biomass production. According to Ficken et al. (2000), Paq values, between 0.4 and 1 characterize submerged and floating aquatic plants, while terrestrial plants yield values lower than 0.1. Indeed, average HI values of ~ 250, together with elevated Paq values support the interpretation that mainly lacustrine primary biomass was produced under medium to high water levels, generating anoxic conditions in B4 depression. TOC values substantially decrease at 160 cm (16.3 cal kyr BP) accompanied by sand maxima. This likely documents an extremely low stand of the water table and the development of a desiccation surface. Increased sand input from the depression margins may take place because of temporary heavy rains during dry phases when the sparse vegetation cover protected less against erosion. Decreasing Paq values between 140 and 120 cm shows higher terrestrial input, likely due to higher rainfall intensities and accelerated erosion at the margins of the depression. Nevertheless, water levels dropped again already at ~ 15.4 cal kyr BP as evidenced by the formation of a swamp around 15.4–15.2 cal kyr BP (120–110 cm; Fig. 4). Decreasing TOC and HI values and absence of pyrite in sediments deposited between 14.4 and 13.6 cal kyr BP indicate accelerated local desiccation, likely correlated with periodic sediment input. Varying sand contents suggest that humid and dry phases alternated, leading to deflation of small particles and organic matter, leaving behind sand in the depression. Intensive deflation and complete desiccation are likely responsible for the hiatus at 70 cm depth (13.6–10.2 kyr BP), coinciding with the transition from Late Glacial to Holocene.

Stable C isotopes are frequently used in paleovegetation reconstructions due to their potential in discerning between C3 and C4 plants (Eshetu 2002; Glaser and Zech 2005). Our bulk δ¹³C values and those of long-chain n-alkanes show average values of −14.3‰ and –25.9‰, respectively. This might show C4 vegetation (Ficken et al. 2002; Glaser and Zech 2005) growing at the margin of the depression under relatively dry conditions during the Late Glacial. However, methanogenesis, assimilation of ¹³C-enriched CO₂ and HCO₃⁻ by water plants due to long-lasting ice cover, especially during the early Late Glacial, and the low CO₂ concentration in the atmosphere (Monnin et al. 2001) might have contributed to such positive δ¹³C values (Conrad et al. 2007). In contrast, modern plants from the surroundings of the lake show values indicating C3 vegetation. This result agrees with a study of Mekonnen et al. (2019), reporting that the modern vegetation in high altitudes of the Bale Mountains comprises only C3 plants.

BC produced by incomplete combustion of plant biomass is ubiquitous in lake sediments and glaciers (Glaser 1998). The weak BC maximum in the lowermost samples (~ 250 cm) presumably reflects BC storage in glacier ice during the LGM, originating from atmospheric input. In contrast, high BC between 16.9 and 16.6 cal kyr BP and between 15.7 and 14.4 cal kyr BP provides evidence for frequent fire events, likely due to increased amounts of combustible biomass under regionally wetter than under dry conditions.

Inorganic geochemistry

After deglaciation, the period between 18.1 and 16.6 cal kyr BP started with the accumulation of electrolyte-poor melting water in depression B4 (see EC in Additional file 1). The presence of pyrite at the bottom of the B4 profile implies a sedimentary environment with mainly anoxic conditions and sufficient organic matter that acts as a reductant and energy source for pyrite formation (Berner 1984). Due to their geochemical stability, lithogenic elements such as Zr, Hf, Nb, and Ti are often used in paleoenvironmental studies as indicators of detrital and allochthonous input (Davies et al. 2015). For instance, Zr and Zr based ratios (Zr/Ti, Zr/Rb, Zr/Al) are often used as proxies for aeolian input of Saharan dust (Jimenez-Espejo et al. 2014; Scheuvens et al. 2013; Moreno et al. 2006, and Hemming 2007), while Si ratios (Ti/Si, Zr/Si) are used as indicators of biogenic silica production during favorable seasons (Lamb et al. 2018; Davies et al. 2015; Brown et al. 2007). Furthermore, also K is used as an effective moisture fluctuation indicator (Foerster et al. 2012). In our B4 profile, lithogenic elements and ratios such as Zr, Hf, Zr/SiO₂, Zr/Rb, and Zr/TiO₂ show minimum values after deglaciation; they slightly increased around 210 and 190 cm (~ 17.2 and 16.9 cal kyr BP), while a continuous increase of these elements and ratios is recorded from 175 to 135 cm (~ 16.6–15.7 cal kyr BP) (Fig. 5). In contrast, CPA values show a weak minimum at ~ 210 and 190 cm and further decrease to < 80 between 175 and 135 cm. These results likely reflect intensified dry northerly winds (Brown et al. 2007; Marshall et al. 2011; Lamb et al. 2018; Jimenez-Espejo et al. 2014), which transported less weathered (low CPA, Na enriched, Fig. 5), allochthonous material in the Sanetti Plateau. The occurrence of gypsum-bearing layers in ~ 210 cm (~ 17.2 cal kyr BP) and between 166 and 152 cm (16.4–16.1 cal kyr BP) show lower water tables and provide evidence for dry events. The latter is further confirmed by increasing values of K₂O (Fig. 5), which agrees with results from Chew Bahir (Foerster et al. 2012). Moreover, increased values of K₂O and Na₂O (Fig. 5) coincide with the high abundance of Feldspar in ~ 135 cm (Additional file 3), highlighting Feldspar containing allochthonous input. The abrupt decrease of most allochthonous elements and ratios (Fig. 5) at about 15.7 cal kyr BP indicates a drastic atmospheric reorganization with an abrupt decrease of dust inputs related to the beginning of the AHP, attributed to the northward migration of the rain belt and the ITCZ (Bastian et al. 2021).

Palynology

The green algae Botryococcus is generally found in tropical freshwater lakes under very cold environmental conditions (Jankovská and Komárek 2000). Therefore, the high occurrence of Botryoccoccus at the bottom of the B4 profile indicates deglaciation and discharge of cold meltwater into the lake. The abundant presence of Botroyoccocus, which is usually characterized by high TOC/N values of 18–36 (Last and Smol 2002; Bittner et al. 2020) and high δ¹³C values (Grice et al. 2001), may have contributed to the TOC/N increase recorded between 250 and 240 cm depth and positive δ¹³C values at the bottom of the profile (Fig. 4). This supports the interpretation of predominant lacustrine TOC production after deglaciation. The decrease of Botryococcus and a parallel increase of Pediastrum also show a relatively stable water body in the investigated depression.

The high percentages of Poaceae and herbs (non-arboreal pollen, NAP) between 18.1 and 17.2 cal kyr BP documents open grass-land vegetation around the Sanetti Plateau. The high number of Artemisia and Chenopodiaceae/Amaranthaceae pollen indicate that the vegetation may have been steppe-like. The abundant presence of Plantago suggests a sparse vegetation cover with less competition shortly after deglaciation. However, the presence of arboreal pollen points to forests down the slopes of the Bale Mountains during the LGM. An increase in Chenopodiaceae/Amaranthaceae and Plantago and a decrease in Artemisia, Pediastrum, Hagenia, Podocarpus, Myrica, Juniperus, and Olea between ~ 17.2 and 16.9 cal kyr BP reflect a reduction of the montane forests due to a ~ 300-year interval with dry and/or cold climate. This interpretation further agrees with the mineralogical results which recorded a dry event between 17.2 and 16.9 cal kyr BP. The increase of arboreal pollen and Pediastrum between about 16.9–16.6 cal kyr BP marks a recovery of the montane forest and wetter local setting at the slopes of the Sanetti Plateau. At around 16.5 cal kyr BP, low values of Pediastrum, Erica, Hagenia, and Botryococcus indicate dry environmental conditions. The decrease in the influx of Erica pollen and subsequent increase of Chenopodeacea/Amaranthaceae around 16.3 cal kyr BP related to the desiccation  phase of the B4 depression recorded by low TOC and high sand input. After this desiccation phase, a trend to locally more humid conditions is indicated by higher Pediastrum values, more frequent fern spores, and Sparganium-type between 155 and 135 cm (16.1 and 15.8 cal kyr BP). This is in agreement with increasing TOC values and it documents that on the SP the H1 stadial is characterized by a short-term desiccation, in contrast to the lowland archives (Fig. 8). Dry environmental conditions are also evidenced from 15.4 to 15.2 cal kyr BP by the decrease in Erica and Hagenia pollen, while tolerant montane taxa such as Podocarpus and Juniperus survived. Nevertheless, as indicated by constant high Pediastrum influx and high Paq values, the B4 depression was partly or periodically flooded. Changes in seasonality and longer dry seasons may have given way to swamp formation. However, pollen types, including typical swamp and water plants (Cyperaceae, Sparganium-type), show no increase. The increasing Plantago-type includes the amphibious species Littorella uniflora, but this swamp species is not known from Africa today (Hoggard et al. 2003). Possibly, the increased Poaceae influx marks the spreading of swamp grasses at the site. A temporary decrease in the influx of arboreal pollen may be related to a short cold event that also decreased the flourishing of grasses and herbs (NAP). The long-time trend in NAP influx reduction can relate to a change in the vegetation from wind-pollinated plant groups (Poaceae, Chenopodiaceae/Amaranthaceae, Plantago-type, Rumex-type) to insect-pollinated taxa (Senecio-type, Caryophyllaceae), leading to an overall reduced NAP pollen input. By contrast, the decrease in Chenopodiaceae/Amaranthaceae and consequent increase of A/C between 15.2 and 14.4 cal kyr BP indicate the most humid conditions regionally, whereas locally the B4 depression started to dry out due to increased silting up. A significant increase in Erica and Podocarpus pollen between 14.4 and 13.6 cal kyr BP substantiates the expansion of the Ericaceous belt and the lowland forests, together with a fundamental change in vegetation composition on the Sanetti Plateau. This is in contrast to the sedimentological results which document progressive local drying out. This discrepancy might be due to the potential of pollen assemblage to provide regional environmental signals, while biogeochemical proxies (Fig. 4), mainly reflect the local conditions of the B4 record.

Fig. 8

A summary figure comparing palynological and biogeochemical results from unit 1 and unit 2 of the B4 sediments with relevant proxies from Lake Tanganyika (Tierney et al. 2008), Lake Challa (Tierney et al. 2011) and Lake Chew Bahir (Foerster et al. 2012) sediments. Light orange color indicate dryer events, orange color indicates a desiccation layer developed during Heinrich Event 1 (H1). Blue color characterizes the African Humid Period (AHP) and other humid episodes while dark green color indicates the event characterized as humid and arid according to pollen and geochemical results, respectively. δD and K values are in inversed scale

Environmental implications and comparison with other records

Deglaciation and climate fluctuations

¹⁴C ages from the lower sediment of depression B4 suggest that the Sanetti Plateau became at least locally, ice-free already at ~ 18.2 cal kyr BP. Surface exposure dating of erratic boulders in N-exposed valleys and on the Sanetti Plateau suggest the onset of deglaciation at 15.2 ± 1.2 cal kyr BP (Groos et al. 2021; Ossendorf et al. 2019). The basal ¹⁴C-age of Garba Guracha sediments (15.9–16.7 cal kyr BP) is almost in agreement with the age of deglaciation estimated from cosmogonic ages (Umer et al. 2007; Tiercelin et al. 2008; Bittner et al. 2020). This discrepancy might be due to the ice cover in the N-exposed trough valley of Lake Garba Guracha that might have been thicker and thus longer lasting than on the Plateau around the B4 profile, which is strongly wind-exposed and has no ice accumulating catchment (Fig. 1). Besides, methodological uncertainties (exposure dating versus radiocarbon analysis) cannot be ruled out. Nevertheless, the deglaciation age of B4 depression seems reliable.

Our palynological and biogeochemical results record climate fluctuations on the Sanetti Plateau since the end of the LGM. Disregarding the initial phase around 18 cal kyr BP, the environmental conditions during unit 1 can be characterized as predominantly lacustrine, correlating with results from Lake Tanganyika, Challa, and Chew Bahir, which also show predominantly humid conditions (Fig. 8). However, between ~ 17.2 and 16.9 cal kyr BP partly fluctuating allochthonous elements (Fig. 5), correlating with increased deflation in the northerly dust source areas, and an increase in Chenopodiaceae/Amaranthaceae and reduction in montane forest indicate a short dry event during this time. Possibly, this event relates to a short weakening of the East-Asian monsoon system as reported by Wang et al. (2001) for Asia around 17 cal kyr BP as a precursor of Heinrich Event 1 (Hemming 2004). Also, Camuera et al. (2021) described such an early H1 phase (refers to HS1b) between 17.2–16.9 cal kyr BP in the Mediterranean. The ~ 300 years of climate deterioration are also weakly documented in sediments of Chew Bahir (Fig. 8) and might be part of the mega-drought postulated for the Afro-Asian monsoon system between 17–16 cal kyr BP (Stager et al. 2011).

We interpret increasing input of allochthonous elements between 16.6 and ca.15.9 cal kyr BP (Fig. 5) as an indicator of the dominance of northerly wind, correlated with progressive ecosystem degradation in the lowlands, and occurring contemporaneously with the North Hemispheric cooling during H1. Interestingly, on the Sanetti Plateau TOC-minima and sand-maxima indicate only short-term intensive desiccation at 16.3 cal kyr BP, and already between ~ 16.1 and 15.7 cal kyr BP TOC and HI increased again, whereas sand contents rapidly decreased (Fig. 4), reflecting that the environmental conditions locally, on the Sanetti Plateau improved, in agreement with the palynological results. This discrepancy between the high altitudes of the Sanetti Plateau and the vast lowlands northerly might reflect that the Bale Mountains, located close to the Indian Ocean, react more rapidly to environmental changes, whereas the lowland ecosystems are much more resilient, due to slow negative vegetation and monsoon feedbacks.

In many studies, arid conditions in tropical Africa between ~ 16.8 and 15.4 cal kyr BP were already described, coinciding with H1 (e.g. Bonnefille and Chalie 2000; Gasse 2000; Talbot and Lærdal 2000; Hemming 2004; Tiercelin et al. 2008; Tierney et al. 2008). Lamb et al. (2007) also mentioned an “episode of shallow water” in the Lake Tana basin between 16.7 and 15.7 cal kyr BP. Based on δD values of higher plant leaf waxes from Lake Challa (Fig. 8), Tierney et al. (2011) concluded that these arid conditions in eastern Africa were due to reduced moisture input from the Indian Ocean. Previous studies described that climatic fluctuations in eastern Africa during the Late Quaternary were related to teleconnections between North Atlantic cooling events, recorded in Greenland ice cores and N-Atlantic sediments, and the weakening of the Indian Summer Monsoon (Tierney et al. 2008; Marshall et al. 2011; Mohtadi et al. 2014; Lamb et al. 2018). It is suggested that during these cooling events, caused by massive iceberg surges and cold meltwater entering the North Atlantic, the ITCZ and the Northern Hemisphere westerlies shifted abruptly southwards and abruptly northward at the end of the cooling, thus controlling deflation of lithogenic elements out of the northern source areas and their input in the surrounding areas (Tierney et al. 2008; Marshall et al. 2009; Lamb et al. 2018). This might explain the high input of allochthonous lithogenic elements (Fig. 5). A similar finding is presented by Brown et al. (2007), reporting elevated inputs of e.g. Zr to Lake Malawi due to intensified northerly winds.

The allochthonous elements decreased abruptly at about 15.7 cal kyr BP, likely correlating with the onset of the AHP, and evoked by rapid northward migration of the rain belt and the ITCZ in relation to precession-driven insolation changes (Bastian et al. 2021). We assume that this resulted in a relatively rapid establishment of a denser vegetation cover and a drastic reduction of dust deflation from northern source areas (Figs. 5 and 8). The timing of this event agrees with the results of Camuera et al. (2021). They concluded that the AHP started in the Mediterranean at 15.7 cal kyr BP after the termination of HS1c. Also, Lamb et al. (2007, 2018), described such an abrupt shift from Lake Tana, and Marshall et al. (2011) dated this event to 15.3 cal kyr BP. In Lake Victoria, the rapid refilling of the basin is dated to 14.5 cal kyr BP (Talbot and Lærdal 2000). One reason for this discrepancy might be dating uncertainties. However, our data from the B4 profile let assume that locally, at the high altitudes of the Sanetti Plateau, the abrupt onset of humidity might have started almost 1000 years before the onset of the Bølling–Allerød warming in the North-Atlantic region (Alley and Clark 1999; Van Raden et al. 2013) and before at least some hundred years earlier than in Lake Tana region, located 630 km northward. In contrast to the sedimentological and geochemical results, our pollen results (Figs. 6 and 7) do not show clear evidence for this abrupt start of the AHP. Within this humid period, water levels in the B4 record did not remain high but lowered already around 15.4 cal kyr BP, and a swamp developed, reflecting probably higher evaporation and less precipitation. A similar finding was reported from Lake Tana, where a papyrus swamp developed between 15.7 and 15.1 cal kyr BP, due to lower lake levels and reduced rainfall (Lamb et al. 2007). Slightly rising allochthonous proxies (e.g. Zr, Nb, Hf, see Fig. 5) likely document progressive local drying-up after about 14.4 cal kyr BP, whereas ascending arboreals indicate regional wetter conditions (Fig. 7). The uppermost part of the unit 2 sediments, deposited between ~ 14.4–13.6 cal kyr BP, is enriched in arboreal pollen, reflecting regional wetting; but fluctuating sand contents and higher input of terrestrial alkanes let assume that drying-up of the B4 depression by advanced periodic sediment input continued. These processes do not allow a reliable correlation of the environmental dynamics affecting the Sanetti Plateau with those documented in sediments of Lake Tanganyika, Challa, and Chew Bahir (Fig. 8); also an unambiguous relationship with climatic phenomena known from the North Atlantic region (e.g. Allerød) cannot be identified in the upper part of unit 2.

Vegetation

The B4 pollen record allows the reconstruction of vegetation changes from 18.1 to 13.6 cal kyr BP. Between 18.1 and 16.8 cal kyr BP, the vegetation is characterized as herb-rich grassland with Chenopodiaceae/Amaranthaceae. Plantago maxima between 17.2–16.9 cal kyr BP are related to dry and/or cold environmental conditions (see Section 4.4). These results are roughly in agreement with Umer et al. (2007), reporting open grassland vegetation at 16 cal kyr BP around Lake Garba Guracha. Our pollen record further shows high Arboreals such as Podocarpus, Hygenia, Erica, and Juniperus at 16.9 cal kyr BP, indicating more favorable conditions, in agreement with the sedimentological data. Increased aridity between 16.6 and 15.7 cal kyr BP, well documented by most biogeochemical proxies and interpreted to correlate with H1, is not clearly revealed in the palynological data. The desiccation surface developed around ~ 16.3 cal kyr BP correlates with decreased contents of Erica pollen, Pediastrum, ferns, and Botryococcus, and elevated Chenopodiaceae/Amaranthaceae ratios, indicating a temporary phase of less favorable, drier environmental conditions. This possibly marks the time of maximum H1 dryness. Similarly, the AHP, recorded by biogeochemical proxies, is not obvious in the pollen results. Our data show that Ericaceae pollen content considerably increased on the Sanetti Plateau from 14.4 to 13.6 cal kyr BP, likely correlating with an increase in temperature, which might be synchronous with the European Meiendorf interstadial starting at 14.4 varve years (Litt et al. 2001). According to pollen results from Lake Garba Guracha and peat deposits, Umer et al. (2007) and Kuzmicheva et al. (2014) assume that Erica vegetation expanded on the Sanetti Plateau above the actual upper limit of the Erica belt, only during the early Holocene beginning at 11.2 cal kyr BP. The dry climate during mid to Late Holocene and natural and/or human-induced fires (Miehe and Miehe 1994) are suggested as major causes for the decline of Erica which at present exists above the upper line of the Erica belt only in form of isolated fragments, mainly restricted to slopes rich in basalt boulders. Furthermore, Miehe and Miehe (1994) assumed that these Erica patches are relics that document an intensive extension of Erica species on the Sanetti Plateau after deglaciation, which were later on destroyed because of the invasion of hunter-gatherers and pastoralists. However, since the upper timberline is temperature-controlled (Körner 2008), we hypothesize these Erica patches on the plateau survive due to better microclimate, generated by dark basalt boulders.

Fire dynamics on the Sanetti Plateau

Our black carbon results (Fig. 4) show that fire was a common phenomenon in the Bale Mountains even during the early Late Glacial. However, pollen data indicate that at this time Erica was not growing around the B4 depression (Fig. 8). Therefore, BC in the lower part of the B4 profile likely originates from other vegetation fires or the Erica belt in lower altitudes. This interpretation is supported by the fact that BC, even in the dry mode H1 sediments, is not clearly increased. Only with the beginning of the AHP around 15.7 cal kyr BP, BC started to increase, correlating with warmer and more humid environmental conditions, and reaching a maximum at about 15 cal kyr BP coinciding with increasing Erica pollen. In the uppermost part of unit 2 (70–80 cm) Erica pollen further increased, again correlating with maximum BC at about 13.6 cal kyr BP.

Up to now, despite the human invasion during 47–31 cal kyr BP, there is no evidence that during the Late Glacial hunter-gatherers were living on the plateau and burned the Erica to facilitate hunting. However, we cannot rule out that fires were triggered by the increase of easily combustible Erica biomass due to improved environmental conditions. For a different time period, but in connection with the fuel-controlled fire process, the charcoal record from Garba Guracha supports a strong correlation between fire occurrence and heathland expansion during the Holocene (Gil-Romera et al. 2019). Elevated BC in the early humid Holocene deposits of the B4 record (not shown here) supports our interpretation that the burning of Erica is mainly controlled by the amount of Erica biomass, being higher under warm and humid climates.

Conclusions

Our high altitude sedimentary archive provides a new older deglaciation age of 18.2 cal kyr BP for the Sanetti Plateau. Biogeochemical results show that the Plateau was sensitive to local, regional and global climate changes. We detected a severe local drought event on the Sanetti Plateau at ~ 16.3 cal kyr BP with complete desiccation of the past B4 lake for some decades, related to H1. Between ~ 16.6 and 15.7 cal kyr BP allochthonous elements like Zr, Hf, Nd, Nb, Na, presumably windblown by dry northerly winds during H1, accumulated increasingly in the B4 record, indicating continuous but delayed degradation of the wind source areas in lower altitudes but without documenting maximum dryness at 16.3 cal kyr BP. This is in contrast to our high altitude B4 archive and reflects the resilience of the lowland ecosystems. The Abrupt change to humid conditions at ~ 15.7 cal kyr BP indicates the onset of AHP in the Bale Mountains some hundred years earlier than in the Lake Tana region but in agreement with the termination of the HS1c phase in the Mediterranean.

The vegetation on the Sanetti Plateau was less sensitive to increased aridity during H1 and also to increased humidity during AHP. Nevertheless, the Erica pollen increased in the B4 sediments at ~ 14.4 cal kyr BP, correlating with a wet and warm regional climate. Despite this increase, it remains open whether the Ericaceous belt expanded to the plateau during that time or during the early Holocene, except for the currently existing isolated patches on boulder-rich steep slopes. Our results indicate that fire incidences mainly coincide with an expansion of the vegetation cover and less with dry periods. Most likely, a warm and humid climate promotes biomass production of Erica, hence increasing the amount of fuel, which burns from time to time. This allows to conclude that biomass and thus fuel availability is important factor controlling fire events in the Bale Mountains.